Permocupedidae
Permocupedidae is a family of Protocoleopteran stem group beetles. They first appeared during the Early Permian, and were one of the dominant groups of beetles during the Middle Permian. They became rare in the Late Permian, with only one species known from the Triassic, Frankencupes ultimus from the Middle Triassic (Anisian) Röt Formation of Germany.[1] They are thought to have been xylophagous (wood eating), which is presumed to be the ancestral ecology of beetles.[2] One monotypic with the species Linicupes yinpinensis, was described in 2014 from late Permian fossils in found in China.[3]
| Permocupedidae Temporal range: | |
|---|---|
 | |
| Fossil and life restoration of Permocupes sojanensis | |
Scientific classification  | |
| Kingdom: | Animalia |
| Phylum: | Arthropoda |
| Class: | Insecta |
| Order: | Coleoptera |
| Suborder: | †Protocoleoptera |
| Superfamily: | †Permocupedoidea |
| Family: | †Permocupedidae Martynov, 1933 |
Taxonomy
| Afrocupes ✝ | 290 Ma
Permian |
| Archicupes ✝ | 272 Ma
Permian |
| Cytocupes ✝ | 272 Ma
Permian |
| Cytocupoides ✝ | 272 Ma
Permian |
| Eocupes ✝ | 272 Ma
Permian |
| Frankencupes ✝ | 247 Ma
Triassic |
| Ichthyocupes ✝ | 272 Ma
Permian |
| Kaltanicupes ✝ | 279 Ma
Permian |
| Kaltanocoleus ✝ | |
| Linicupes ✝ | 265 Ma
Permian |
| Maricoleus ✝ | 279 Ma
Permian |
| Permocupes ✝ | 272 Ma
Permian |
| Protocupes ✝ | |
| Protocupoides ✝ | 279 Ma
Permian |
| Tatarocupes ✝ | 265 Ma
Permian |
| Uralocupes ✝ | 272 Ma
Permian |
References
- Ponomarenko, Alexandr G.; Bashkuev, Alexei S. (December 2018). "First Triassic record of the beetle family Permocupedidae (Insecta: Coleoptera): a peculiar example of a Lazarus taxon". PalZ. 92 (4): 587–591. doi:10.1007/s12542-018-0426-4. ISSN 0031-0220.
- Zhao, Xianye; Yu, Yilun; Clapham, Matthew E; Yan, Evgeny; Chen, Jun; Jarzembowski, Edmund A; Zhao, Xiangdong; Wang, Bo (2021-11-08). Perry, George H; Fikacek, Martin (eds.). "Early evolution of beetles regulated by the end-Permian deforestation". eLife. 10: e72692. doi:10.7554/eLife.72692. ISSN 2050-084X.
- A. G. Ponomarenko, E. V. Yan and Di-ying Huang (2014). "New beetles (Coleoptera) from the terminal Middle Permian of China". Paleontological Journal 48 (2): 191–200. doi:10.1134/S0031030114010109.
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