Tshekardocoleidae

Tshekardocoleidae is an exinct family of beetles, known from the Permian. Like other primitive beetles, they are thought to have been xylophagous.[1] The oldest known beetle, Coleopsis, was originally assigned to this family, but is now assigned to its own family Coleopseidae.[2] They repesent some of the earliest known beetles, first appearing during the Cisuralian, before becoming extinct at the beginning of the Guadalupian.[1] A claimed Jurassic record is doubtful.[2]

Tshekardocoleidae
Temporal range:
Fossil and life restoration of Moravocoleus permianus
Scientific classification
Kingdom: Animalia
Phylum: Arthropoda
Class: Insecta
Order: Coleoptera
Suborder: †Protocoleoptera
Family: †Tshekardocoleidae
Rohdendorf, 1944
Genera

See text

Taxonomy

After[2]

  • Avocoleus Ponomarenko, 1969 Type species: Moravocoleus fractus Kukalová, 1969, by original designation. Two species. Czech Republic: Moravia (Obora), Permian, Cisuralian, Sakmarian, −295.0–290.1 Ma.
  • Boscoleus Kukalová, 1969 Type species: Boscoleus blandus Kukalová, 1969, by monotypy. One species. Czech Republic: Moravia (Obora), Permian, Cisuralian, Sakmarian, −295.0–290.1 Ma.
  • Eocoleus Kukalová, 1969 . Type species: Eocoleus scaber Kukalová, 1969, by monotypy. One species. Czech Republic: Moravia (Obora), Permian, Cisuralian, Sakmarian, −295.0–290.1 Ma.
  • Moravocoleus Kukalová, 1969 . Type species: Moravocoleus permianus Kukalová, 1969 by original designation. One species. Czech Republic: Moravia (Obora), Permian, Cisuralian, Sakmarian, −295.0–290.1 Ma.
  • Permocoleus Lubkin et Engel, 2005 . Type species: Permocoleus wellingtoniensis Lubkin et Engel, 2005, by monotypy. One species. USA: Oklahoma, Noble County (Midco Locality and Tasch Nobel V, Bed 7), Permian, Cisuralian, Sakmarian-Artinskian −290.1–279.5 Ma.
  • Prosperocoleus Kukalová, 1969 . Type species: Prosperocoleus prosperus Kukalová, 1969, by monotypy. One species.Czech Republic: Moravia (Obora), Permian, Cisuralian, Sakmarian, −295.0–290.1 Ma.
  • Retelytron Kukalová, 1965 . Type species: Retelytron conopeum Kukalová, 1965, by monotypy. One species. Czech Republic: Moravia (Obora), Permian, Cisuralian, Sakmarian, −295.0–290.1 Ma.
  • Sylvacoleodes Ponomarenko, 1963 . Type species: Sylvacoleodes admirandus Ponomarenko, 1963, by monotypy. One species. Russia: Perm’ Region, Suksun (Tshekarda, Sylva River), Permian, Cisuralian, Artinskian-Kungurian, −279.5–272.5 Ma.
  • Sylvacoleus Ponomarenko, 1963 . Type species: Sylvacoleus richteri Ponomarenko, 1963, by original designation. Two species. Also includes: Sylvacoleus sharovi Ponomarenko, 1963. Russia: Perm’ Region, Suksun (Tshekarda, Sylva River), Permian, Cisuralian, Artinskian-Kungurian, −279.5–272.5 Ma.
  • Tshekardocoleus Rohdendorf, 1944 . Type species: Tshekardocoleus magnus Rohdendorf, 1944, by monotypy. Two species. Also includes: Tshekardocoleus minor Ponomarenko, 1963 . Russia: Perm’ Region, Suksun (Tshekarda, Sylva River), Permian, Cisuralian, Artinskian-Kungurian, −279.5–272.5 Ma.
  • Umoricoleus Kukalová, 1969 . Type species: Umoricoleus perplex Kukalová, 1969, by monotypy. One species. Czech Republic: Moravia (Obora), Permian, Cisuralian, Sakmarian, −295.0290.1 Ma.
  • Votocoleus Kukalová, 1969 . Type species: Votocoleus submissus Kukalová, 1969, by monotypy. One species. Czech Republic: Moravia (Obora), Permian, Cisuralian, Sakmarian, −295.0–290.1 Ma.

References
  1. Zhao, Xianye; Yu, Yilun; Clapham, Matthew E; Yan, Evgeny; Chen, Jun; Jarzembowski, Edmund A; Zhao, Xiangdong; Wang, Bo (2021-11-08). Perry, George H; Fikacek, Martin (eds.). "Early evolution of beetles regulated by the end-Permian deforestation". eLife. 10: e72692. doi:10.7554/eLife.72692. ISSN 2050-084X.
  2. Kirejtshuk, Alexander G. (2020-02-17). "Taxonomic Review of Fossil Coleopterous Families (Insecta, Coleoptera). Suborder Archostemata: Superfamilies Coleopseoidea and Cupedoidea". Geosciences. 10 (2): 73. Bibcode:2020Geosc..10...73K. doi:10.3390/geosciences10020073. ISSN 2076-3263.
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