Crocodylus falconensis

Crocodylus falconensis is an extinct species of crocodile known from the early Pliocene of the lower part of the Vergel Member of the San Gregorio Formation of Venezuela. C. falconensis was named in 2013 and it is thought to be the basalmost known Neotropical crocodile species. The cladogram below follows the topology from Scheyer et al. (2013).[1]

Mecistops cataphractus

 Crocodylus 

C. palaeindicus

 Indopacific 

C. palustris

C. siamensis

C. porosus

C. mindorensis

C. johnsoni

C. novaeguineae

 AfricaNew World 

C. niloticus

 New World 

C. falconensis

C. moreletii

C. rhombifer

C. acutus

C. intermedius

Crocodylus falconensis
Scientific classification
Kingdom: Animalia
Phylum: Chordata
Class: Reptilia
Order: Crocodilia
Family: Crocodylidae
Genus: Crocodylus
Species:
C. falconensis
Binomial name
Crocodylus falconensis
Scheyer et al., 2013

A 2018 tip dating study by Lee & Yates simultaneously using morphological, molecular (DNA sequencing), and stratigraphic (fossil age) data established the inter-relationships within Crocodylidae.[2] In 2021, Hekkala et al. were able to use paleogenomics, extracting DNA from the extinct Voay, to better establish the relationships within Crocodylidae, including the subfamilies Crocodylinae and Osteolaeminae.[3]

The below cladogram shows the results of the latest study, with C. falconensis at the base of an Africa+Americas clade of crocodiles within the genus Crocodylus:

Crocodylidae
Osteolaeminae

Mecistops cataphractus West African slender-snouted crocodile

Euthecodon

Brochuchus

Rimasuchus

Osteolaemus osborni Congo dwarf crocodile

Osteolaemus tetraspis Dwarf crocodile

Crocodylinae

Voay

Crocodylus

Crocodylus anthropophagus

Crocodylus thorbjarnarsoni

Crocodylus palaeindicus

Crocodylus Tirari Desert

Asia+Australia

Crocodylus johnstoni Freshwater crocodile

Crocodylus novaeguineae New Guinea crocodile

Crocodylus mindorensis Philippine crocodile

Crocodylus porosus Saltwater crocodile

Crocodylus siamensis Siamese crocodile

Crocodylus palustris Mugger crocodile

Africa+New World

Crocodylus checchiai

Crocodylus falconensis

Crocodylus suchus West African crocodile

Crocodylus niloticus Nile crocodile

New World

Crocodylus moreletii Morelet's crocodile

Crocodylus rhombifer Cuban crocodile

Crocodylus intermedius Orinoco crocodile

Crocodylus acutus American crocodile

(crown group)

References

  1. Scheyer, T. M.; Aguilera, O. A.; Delfino, M.; Fortier, D. C.; Carlini, A. A.; Sánchez, R.; Carrillo-Briceño, J. D.; Quiroz, L.; Sánchez-Villagra, M. R. (2013). "Crocodylian diversity peak and extinction in the late Cenozoic of the northern Neotropics" (PDF). Nature Communications. 4: 1907. doi:10.1038/ncomms2940. PMID 23695701.
  2. Michael S. Y. Lee; Adam M. Yates (27 June 2018). "Tip-dating and homoplasy: reconciling the shallow molecular divergences of modern gharials with their long fossil". Proceedings of the Royal Society B. 285 (1881). doi:10.1098/rspb.2018.1071.
  3. Hekkala, E.; Gatesy, J.; Narechania, A.; Meredith, R.; Russello, M.; Aardema, M. L.; Jensen, E.; Montanari, S.; Brochu, C.; Norell, M.; Amato, G. (2021-04-27). "Paleogenomics illuminates the evolutionary history of the extinct Holocene "horned" crocodile of Madagascar, Voay robustus". Communications Biology. 4 (1): 1–11. doi:10.1038/s42003-021-02017-0. ISSN 2399-3642.
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